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Paul Opler

Dr. Opler

Entomology

Professor
018 Hartshorn Building

C.P Gillette Museum

Paul.Opler@ColoState.edu
970-667-8448 office
970-491-0490 lab

Research Interests:

My research involves a number of separate projects that relate to the systematics, genetics, biological diversity, distribution, and ecology of several groups of insects, primarily Lepidoptera and Apoidea (Hymenoptera). Currently, I am actively monitoring the flowering plants visited by Colorado and Wyoming bees for nectar and pollen as well as the systematics of selected butterfly genera and species groups including both genetic and morphological characters. I am working on development that benefits the C.P. Gillette Museum of Arthropod Diversity.

Selected Publications:

  1. Taxonomic and distributional studies on the western components of the Apodemia mormo complex (Riodinidae). Jour. Lepid. Soc. 15: 145-171. With J. A. Powell.

 

  1. Some notes on Callophrys (Mitoura) johnsoni in California. Jour. Lepid. Soc. 16: 193-194.

 

  1. Studies on the Nearctic Euchloe. Part 3. Complete synonymical treatment. Jour. Res. Lepid. 5: 185-190.

 

  1. Studies on the Nearctic Euchloe. Part 4. Type data and type locality restrictions. Jour. Res. Lepid. 5: 190-195.

 

  1. A possible record for the occurrence of Callophrys (Xamia) xami in California. Jour. Lepid. Soc. 20: 212.

 

  1. Biological observations on Callophrys viridis (Lycaenidae). Jour. Lepid. Soc. 2:113-114. With R. M. Brown.

 

  1. New host plant records for Anthocaris (Pieridae). Jour. Lepid. Soc. 21: 212.

 

  1. Studies on the Nearctic Euchloe. Part I. Introduction. Jour. Res. Lepid. 5: 39-40.

 

  1. Studies on the Nearctic Euchloe. Part 2. Chronological review of the literature and bibliography. Jour. Res. Lepid. 5: 41-50.

 

  1. A distributional analysis of the butterflies of Contra Costa County, California. Jour.

Lepid. Soc. 22:89-107. With R. L. Langston.

 

  1. Unusual numbers of Hepialus sequoiolus Behrens in Sonoma County. Pan-Pac. Ent. 44:

 

  1. Myrmecophily reported for Icaricia acmon and Philotes enoptes bayensis (Lycaenidae,

Lepidoptera). Pan-Pac. Ent. 44: 79-80.

 

  1. Eurema salome in Texas. Jour. Lepid. Soc. 23: 188.

 

  1. Two new species of Caloptilia associated with Rhus in California (Gracilariidae:

Lepidoptera). Pan-Pac. Ent. 44: 79-80.

 

  1. Book review: Higher classification, phylogeny and zoogeography of the Satyridae

(Lepidoptera), by L. D. Miller. Jour. Lepid. Soc. 23: 276~277.

 

  1. Book review: The North American Aegeriidae (Lepidoptera): a revision based on late

instar larvae, by M. R. McKay. Jour. Lepid. Soc. 23:126.

 

  1. Studies on the Nearctic Euchloe. Part 5. Distribution. Jour. Res. Lepidoptera.

 

  1. The types of lycaenid butterflies described by William Henry Edwards. Part I. Theclinae.

Trans. Amer. Ent. Soc. 96:19-77. With F. M. Brown.

 

  1. Studies on the Nearctic Euchloe. Part 6. Systematics of adults. Jour. Res. Lepid. 8:153-

168.

 

  1. Seasonal distribution of Macrolepidoptera in Santa Clara County, California. Jour. Res.

Lepid. 9: 75-88. With J. S. Buckett.

 

  1. Biological and systematic considerations on the “Emarginana-group” of the genus

Epinotia (Torticidae). Jour. Lepid. Soc. 25:115-123.

 

  1. Seven new leaf-miners associated with Quercus agrifolia (Heliozelidae, Gracilariidae).

Jour. Lepid. Soc. 25:194-211.

 

  1. Mass movement of Tarnetrum corruptum (Odonata: Libellulidae). Pan-Pac. Ent. 47: 223.

 

  1. Stigmatic exudates and pollination. Pp. 47-60 in: Pollination and dispersal symposium

offered to Dr. L. van Pijl on his 70th birthday. N. B. M. Brantjes and J. F Linskens, eds. Dept.

Botany, Univ. Nijmegan, Netherlands. With H. G. and I. Baker.

 

  1. Fossil lepidopterous leaf mines demonstrate the age of some insect-plant relationships.

Science 179: 1321-1323.

 

  1. Lepidoptera in Costa Rica. Lepid. News, 1973: 6.

 

  1. Biology, ecology, and host specificity of Microlepidoptera associated with Quercus

agrifolia (Fagaceae). Univ. California Publ. Entomol. 75: 1-83.

 

  1. Distributional relationships of xerophilic insects in the San Joaquin Valley, California.

Southwest. Nat. 18: 305-316.

 

  1. Oaks as evolutionary islands for leaf-mining insects. Amer. Sci. 62: 67-73.

 

  1. Tropical plant phenology: applications for studies in community ecology. Pp.287-296 in:
  2. Lieth (ed.) Phenology and Seasonality Modeling. Springer-Verlag, Berlin. With G. W.

Frankie and H. G. Baker.

 

  1. Bat activity and pollination of Bauhinia pauletia: plant-pollinator coevolution. Ecology

55: 412-419. With E. R. Heithaus and H. G. Baker.

 

  1. Comparative phenological studies of trees in tropical wet and dry forests in the lowlands

of Costa Rica. Jour. Ecol. 62: 881-919. With G. W. Frankie and H. G. Baker.

 

  1. Studies on Nearctic Euchloe. Part 7. Comparative life histories, hosts, and the

morphology of Immature stages. Jour. Res. Lepid. 13:1-20.

 

  1. Population biology and adult behavior of Lycaena xanthoides (Lycaenidae). Jour. Lepid.

Soc. 29: 6366. With J. A. Scott.   

 

  1. Subfamily Lycaeninae. In: W H. Howe, ed. The Butterflies of North America. Doubleday

and Co., Garden City, N.Y.

 

  1. Dioecism in tropical forest trees. Evolution 29: 167-179. With K. S. Bawa.

 

  1. Foraging pattern and resource utilization in seven species of bats in a seasonal tropical

forest. Ecology 56: 841-854. With E. R. Heithaus and T. H. Fleming.

 

  1. Reproductive biology of some Costa Rican Cordia species (Boraginaceae). Biotropica 7:

234-247. With H. G. Baker and G. W. Frankie.

 

  1. Conservation of Lepidoptera. Lepid. News 1975: 7.

 

  1. Rainfall as a factor in the release, timing, and synchronization of anthesis by tropical

trees and shrubs. Jour. Biogeogr. 3: 231-236. With G. W. Frankie and H. G. Baker.

 

  1. Foraging behavior of solitary bees: implications for outcrossing of a Neotropical tree

species. Jour. Ecology 64:1049-1057. With G. W. Frankie and K. S. Bawa.

 

  1. Book review: Ecology of Plants in the Tropics by D. H. Janzen. BioSci. 26: 640.

 

  1. The parade of passing species: extinctions past and present. Science Teacher 43: 30-34.

 

  1. Island biogeography and conservation: strategy and limitations. Science 193: 1030-1032.

With R.F. Whitcomb, J. Lynch and C. Robbins.

 

  1. Spatial relationships between staminate and pistillate plants of dioecious tropical forest

trees. Evolution 31: 64-68. With K. S. Bawa.

 

  1. Interaction of plant life history components with arboreal herbivores. Pp.23-31 in: E.

Montgomery, ed. Ecology of Arboreal Folivores. Symp. Nat’l. Zool. Park.

 

  1. Recovery of tropical lowland forest ecosystems. Pp. 379-421 in: J. Cairns, ed. Recovery of

Damaged Ecosystems. Univ. Virginia Press, Charlottesville. With H. G. Baker and G. W.

Frankie.

 

  1. A review of North American Rhodophaea (Phycitinae, Pyralidae), with description of six

new species. Jour. Lepid. Soc. 31: 41-56.

 

  1. Book review: Libro Rojo de los Lepidopteros Ibericos by M. Viedina and M. Gomez

Bustillo. Jour. Lepid. Soc. 31(4):284.

 

  1. Book review: Legion of Night: The Underwing Moths by T D. Sargent. Bull. Entomol.

Soc. Amer. 23(2):149-150.

 

  1. Book review: Moths of Southern Africa by E. Pinhey. Bull. Entomol. Soc. Amer.

23(2):150-151.   

 

  1. Why are pistillate inflorescences eaten less than staminate inflorescences of Simarouba

glauca? Evolution 32: 673-676.

 

  1. Insects of American chestnut: possible importance and conservation concern. Pp. 83-85

in: R. MacDonald, ed. Proc. American chestnut symp. Univ. West Virginia, Morgantown.

 

  1. A comparison of the amino acid complements of floral and extrafloral nectars. Bot. Gaz.

139: 322-332. With H. G. and I. Baker.

 

  1. Freshwater and terrestrial arthropods. Pp. 171-183. In: D. Linzey, ed. Endangered and threatened species of Virginia. Univ. Virginia Press, Charlottesville.

 

  1. Sex ratios in tropical forest trees. Evolution 32: 812-821. With K. S. Bawa.

 

  1. Plant reproductive characteristics during secondary succession in Neotropical lowland

forest ecosystems. Biotropica Suppl. 40-46. With H. G. Baker and G. W. Frankie.

 

  1. Taxonomic isolation and the accumulation of herbivorous insects: a comparison of

introduced and native trees. Ecol. Ent. 5:205-211. With E. Connor et al.

 

  1. Comparative phenological studies of treelet and shrub species in tropical wet and dry

forests in the lowlands of Costa Rica. J. Ecol. 68:167-188. With G. W. Frankie and H. G. Baker.

 

  1. Polymorphic mimicry of polistine wasps by a Neotropical neuropteran. Biotropica

13:165-176.

 

  1. Insect conservation. Annu. Rev. Ent. 26: 233-258. With R. M. Pyle and M. Bentzien.

 

  1. The leaf-mining moths of the genus Cameraria associated with Fagaceae in California.

Smithsonian Contr. Zool. 333:1-58. With D. R. Davis.

 

  1. Management of prairie habitats for insect conservation. Jour. Nat. Areas Assoc. 1(4): 3-6.

 

  1. Reproductive biology of plants in tropical forests. Pp.183-223 in: H. Lieth and F. Golley,

eds. Tropical rain forest ecosystems: structure and function. Ecosystems of the World, 14A.

Elsevier. With H. G. Baker et al.

 

  1. Fossil leaf-mines of Bucculatrix (Lyonetiidae) on Zelkova (Ulmaceae) from Florissant,

Colorado. Jour. Lepid. Soc. 36: 145-147.

 

  1. Butterfly species accounts. pp. 47-54. In: H. Boyd, Ed. Invertebrates. New Jersey’s

Endangered and Threatened Plants and Animals. Center for Environmental Research,

Pomona, NJ.

 

  1. A vegetative key to the Piperaceae and Melastomataceae of Finca La Selva, Costa Rica.

Brenesia 21: 261-268. With S. Waltz.

  1. Characteristics and organization of the large bee pollination system in the Costa Rica

Dry Forest. pp. 411-447. In: C. E. Jones and R. J. Little, eds. Handbook of Experimental

Pollination Biology. Van Nortstrand and Co. New York. With G. W. Frankie, W. Haber and K.

Bawa.

 

  1. Nectar production in a tropical ecosystem. Pp. 30-79. In: B. Bentley and T. Elias, eds.

Biology of Nectaries. Columbia Univ. Press. New York.

 

  1. Natural history of seven hairstreaks in coastal North Carolina. Jour. Lepid. Soc. 37: 97-
  2. With S. Gifford.

 

  1. Studies on the Nearctic Euchloe. Part 8. Euchloe olympia. Ann. Camegie Mus. 52: 41-54.

With H. Clench.

 

  1. Butterfly species need monitoring. Audubon Naturalist 9(5): 1, 7.

 

  1. Climaciella brunnea. pp.711-712. In: D. H. Janzen, ed. Costa Rican Natural History.

Univ. Chicago Press, Chicago.

 

  1. Cordia alliodora. pp.219-221. in: D. H. Janzen, ed. Costa Rican Natural History. Univ.

Chicago Press, Chicago. With D. H. Janzen.

 

  1. Centris aethyctera and C. fuscata. pp.707-708. In: D. H. Janzen, ed. Costa Rican Natural

History, Univ. Chicago Press, Chicago. With G. Frankie and B. Vinson.

 

  1. Butterflies East of the Great Plains: an Illustrated Natural History. Johns Hopkins Univ.

Press, Baltimore. 294 pp., 54 pls. With G. Krizek.

 

  1. Chapter 2. Invertebrates. pp. 79-165. In: H. Genoways and E. Brenner, eds.

Species of Special Concern in Pennsylvania. Carnegie Museum Natural History Spec. Publ. 11.

With D. Schweitzer and M. Bentzien.

 

  1. Endangered and threatened insects of Maryland. pp. 267-272. In: Norden, Forester, and

Fenwick, eds. Threatened and Endangered Plants and Animals of Maryland. Maryland

Natural Heritage Program Spec. Publ. 84-I, Annapolis, Md.

 

  1. Butterfly counts – 1982 & 1983. Suppl. to Atala, Xerces Soc. Co-ed. with J. Powell.

 

  1. Book review: The Life Histories of Butterflies of Japan. Volume 1. Papilionidae, Pieridae,

Danaidae, by Fukada, Hama, Kuzuya, Takahashi, Tanaka, Tanaka, Wakabayashi, and

Watanabe. Hoikusha Publ. Co., Osaka, Japan.

 

  1. Butterfly counts – 1984. Suppl. to Atala, Xerces Soc. Co-edited with J. Powell.

 

  1. Butterfly-watching tips, how to photograph butterflies. pp. 865-872. In: R. M. Pyle. The

Audubon Society Field Guide to North American Butterflies. Second Printing. A.A. Knopf,

New York.

  1. Distributional patterns of forest insects. P. 17 in Proceedings thirty-sixth annual western

forest insect work conference. Canadian Forestry Service, Edmonton, Alta.

 

  1. Lange’s Metalmark Butterfly (Apodemia mormo langei). In: R. L. DiSilvestro, Ed.

Audubon Wildlife Report – 1986. Audubon Society, New York. With L Robinson.

 

  1. Book review: The preservation of species: the value of biological diversity. by B. G.

Norton. Wildlife Review 202.

 

  1. Book review: Eskimo curlew–a vanishing species? by J. Gallop et al. Wildlife Review

 

  1. Book review: Insect Conservation, an Australian Perspective, by T. R. New. Series

Entomoligica, Vol. 32. W. Junk, Dordrecht, Netherlands, 184 pp. Biol. Review.

 

  1. Butterfly counts – 1985. Suppl. to Atala Vol.14, Xerces Soc. Co-edited with J. Powell.

 

  1. Attracting butterflies to the eastern Colorado yard and garden. Service in Action.

Colorado State University Cooperative Extension. No.5.504. With W. Cranshaw.

 

  1. A new Euchloe (Pieridae) from northwestern Mexico. Jour. Lepid. Soc. 43(3):188-190.

 

  1. An atlas of the butterflies and skippers of the upper Dogue Creek drainage, Fairfax

County, Virginia, 1978-1982. Atala 12(2):21-44.

 

  1. Notes on the rare skipper, Problema bulenta. Jour. Res. Lepid. 25(2):146-148. With G. O.

Krizek.

 

  1. Book review: The Butterflies of North America by J. A. Scott. Naturalist Review. Winter
  2. Page 6.

 

  1. Endangered species status update. News of the Lepid. Soc. Issue 1.

 

  1. Book review: Foundations for a National Biological Survey, ed. by K. C.

Kim and L. Knutson. Bull. Entomol. Soc. Amer. 33(4):265.

 

  1. Butterfly counts – 1986. Suppl. to Atala Vol. 15, Xerces Soc. Co-edited with J. A. Powell.

 

  1. Book review: Auks: an Ornithologist’s Guide by R. Freethy. Wildlife Review. 207:657.

 

  1. North American invertebrate surveys in North America are necessary. Wings (Xerces

Society) 12: 8-10.

 

  1. Butterfly counts – 1987. Suppl. to Atala Vol.16, Xerces Soc. Co-edited with J. W. Brown.

 

  1. Book review: The Science of Scarcity and Diversity, ed. by M. Soulé. Wildlife Review.

208: 372.

 

  1. Book review: Butterflies of North America, by J. A. Scott. Jour. Res. Lepid. 26: 267-270.
  2. Butterflies of the American West–a Coloring Album. Roberts Rinehart. Boulder, Colo.

(with S. Strawn).

 

  1. Book review: The Birds of Canada by W.E. Godfrey. Wildlife Review. 207: 658.

 

  1. Book review: Amazonian Rain Forests, Ecosystem Disturbance and Recovery, ed. by C.
  2. Jordan. Wildlife Review. 211:612.

 

  1. Book review: Birds of the Middle East and North Africa, by Hollom et al. Wildlife

Review 211: 612.

 

  1. Seasonal polyphenisin and ecology of Eurema daira (Pieridae). Pages 515-533 in: The

Evolutionary Ecology of Plants, ed. by J. Bock and Y. Linhart.

 

  1. Book review: Butterflies of Costa Rica, by P. DeVries. Evolution. With G. Austin.

 

  1. Butterflies of Eastern North America, a Coloring Album and Activity Book. Roberts

Rinehart Publishing Co., Boulder, Colo. With S. Strawn.

 

  1. Bulletin of the ESA: Looking ahead. Bull. Entomol. Soc. Amei; 35: 4.

 

  1. Book review: Conservation and Management of Rare and Endangered Plants, ed. by T.
  2. Elias. Wildlife Review 212: 284.

.

  1. Book review: Extinct Birds, by Errol Fuller. Wildlife Review 212: 366.

 

  1. Book review: The Book of Indian Birds, by Salim Ali. Wildlife Review 212: 411.

 

  1. Waterpumping in Lamproptera meges (Papilionidae). Jour. Res. Lepid. 27:143. With

George Krizek,

 

  1. Attracting butterflies to the eastern Colorado garden. The Green Thumb 46(l):24-28.

With Whitney Cranshaw.

 

  1. Fourth of July Butterfly Counts-1988 Report. Xerces Society, Portland, Ore. Co-edited

with J. W. Brown.

 

  1. Patterns of butterfly species density in peninsular Florida. Jour. Biogeogr. 17:615-622.

With J. W. Brown.

 

  1. Fourth of July Butterfly Counts-1989 Report. Xerces Society, Portland, Ore. 47 pp. Coedited with J. W. Brown.

 

  1. Book review: A Guide to the Birds of Costa Rica, by E G. Stiles and A. F. Skutch.

Wildlife Review 218:298.

 

  1. Book Review: Minnesota’s Endangered Flora and Fauna, by B. Coffin and L.

Pfanmuller. Wildlife Review 219:306.

  1. North American problems and perspectives in insect conservation. Pp.9-32 in N.M.

Collins and J. A. Thomas, eds. The conservation of insects and their habitats. Royal

Entomological Society of London. Academic Press, London.

 

  1. Seasonality of climbers: a review and example from Costa Rican dry forest. Pp.369-383

in Biology of Vines, ed. by H. Mooney, F. Putz, and S. Bullock. Cambridge Univ. Press. With H.

  1. Baker and G. W. Frankie.

 

  1. Habitat enhancement for urban invertebrates. Pp. 80-87 in: E. A. Webb and S. Q. Foster,

eds. Perspectives in urban ecology. Denver Museum of Natural History and Thorne Ecological

Institute, Boulder, Colo.

 

  1. Fourth of July Butterfly Counts-1990 Report. Xerces Society, Portland, Ore. 49 pp.

 

  1. Book Review: Invertebrates, by R. C. Brusca and G. J. Brusca. Wildlife Review 222:282.

 

  1. Book Review: Immature Insects, by F. W. Stehr. BioScience. 42(2):132-133.

 

  1. Field Guide to Butterflies of Eastern North America. Roger Tory Peterson Series.

Houghton-Mifflin Co., Boston. 396 pp.

 

  1. Biodiversity and occurrence of prairie Lepidoptera (butterflies and moths). Pages n.p. in:

Proceedings of the nongame wildlife workshop. U.S. Fish and Wildlife Service, Region 6,

Golden, Colo.

 

  1. Fourth of July butterfly counts (1991). Xerces Society, Portland, Ore. 63 pp. Co-edited

with A. B. Swengel.

 

  1. Foreword. In: Iftner, Shuey, and Calhoun, eds. Butterflies and skippers of Ohio. Ohio

Biological Survey, Columbus, Ohio.

 

  1. Atlas of western USA butterflies including adjacent parts of Canada and Mexico. Ray E.

Stanford and Paul A. Opler, Denver, Colo. 285 pp. With R. E. Stanford.

 

  1. New additions to endangered species list. News of the Lepidopterists’ Society. 1993(2):52.

 

  1. Fourth of July butterfly counts (1992). Xerces Society, Portland, Ore. 75 pp. Co-edited

with Ann B. Swengel.

 

  1. Butterflies East and West–a Book to Color. Roberts Rinehart Publ. Co., Niwot, Colo. 80
  2. With S. Strawn.

 

  1. Moths of western North America. 1. Distribution of Saturniidae of western North

America. Contributions of the C. P. Gillette Museum Arthrropod Diversity, Colorado State

University, Fort Collins, Colo. With R. S. Peigler.

 

  1. Newcastle Disease virus found in double-crested cormorants in the United States.

Research Information Bulletin (U.S. Fish and Wildlife Service, Division of Research). 93-13.

  1. What to do about species names: a bibliographic guide. CBE views 16:79-80.
  2. Book Review: Butterflies of the Florissant Region of Colorado, by B. Drummond, T.

Emmel, and M. Minno. Journal of the Lepidopterists’ Society. Vol. 49.

 

  1. The U.S. Endangered Species Act: Conservation and research for aquatic insects.

Journal of Aquatic Conservation 3:289-291.

 

  1. Systematic Compendium of Arthropods and Annelid Worms of Old-growth and Late

Successional Conifer Forests, Mature Riparian Woods, and of Coarse Woody Debris

Associated Arthropods within the Range of the Northern Spotted Owl (Strix occidentalis

caurina). (co-compiler, Jack D. Lattin.) Draft supplemental environmental impact statement on

management of habitat for late-successional and old-growth forest related species within the

range of the northern spotted owl.(appendix). U.S. Forest Service, U.S. Bureau of Land

Management, U.S. Fish and Wildlife Service et al. [Washington, D.C.]

 

  1. Fourth of July butterfly counts (1993). North American Butterfly Association, Baraboo,

Wisc. 55 pp. Co-edited with Ann B. Swengel.

 

  1. Book Review: Butterflies through binoculars, by J. Glassberg. Journal of the New York

Entomological Society.

 

  1. Moths of western North America. 4. Distribution of Oecophoridae of Western North

America. Contributions of the C. P Gillette Insect Biodiversity Museum, Colorado State

University, Fort Collins. With J. A. Powell.

 

  1. First Guide to Butterflies and Moths. Roger Tory Peterson series. Houghton-Mifflin Co.,

Boston, Mass. 128 pp.

 

  1. Conservation of butterfly diversity in North America. Chapter 22. pp. 316-324. In: A. S.

Pullin, Ed. Ecology and Conservation of Butterflies. Chapman and Hall, London.

 

  1. Recovery Plan. Uncompahgre Fritillary Butterfly. U.S. Fish and Wildlife Service,

Denver, Colo. Co-authored with recovery team.

 

  1. Uncompahgre fritillary butterfly demographics: response to Britten et al.. Conservation

Biology 8 (1994), pp. 1156-1157. (A.L. Seidl, first author)

 

  1. Western Palearctic butterflies in photography. Holarctic Lepidoptera 1:7-61. Coauthored

with G. O. Krizek and L. Havel.

 

  1. Presidential address: studying butterflies by photography. Journal of the Lepidopterists’

Society 50.

 

  1. Butterfly diversity and a preliminary comparison with bird and mammal diversity. In:

Wilson, D., ed. Symposium on Biological Diversity. National Academy of Science, Washington

D.C. Co-authored with R. K. Robbins.

 

  1. Conservation and management of butterfly diversity in North America. Pp. 316-324 in: A.S. Pullin, ed. Ecology and Conservation of Butterflies. Chapman and Hall, London.
  2. Moths of western North America. 3. Distribution of Arctiidae of western North America.

Contributions of the C. P. Gillette Insect Biodiversity Museum, Colorado State University, Fort

Collins. with D. C. Ferguson, J. P Donahue, and M. J. Smith.

 

  1. Species richness of western butterflies and moths. in: LaRoe, E. T., ed.

Our Living Resources. National Biological Survey, Washington, D.C.

 

  1. Fourth of July butterfly counts (1994). North American Butterfly Association, Baraboo,

Wisc. (Co-edited with Ann B. Swengel).

 

  1. Fourth of July butterfly counts (1995). North American Butterfly Association, Baraboo,

Wisc. (Co-edited with Ann B. Swengel).

 

  1. Audubon Society’s butterfly gardening (videotape). Mastervision, New York, N.Y. (coauthored with J.A. Ebner)

 

  1. Audubon Society’s butterflies for beginners (videotape). Mastervision, New York, N.Y.

(co-authored with J.A. Ebner).

 

  1. Fourth of July butterfly counts (1996). North American Butterfly Association, Baraboo,

Wisc. (Co-edited with Ann B. Swengel).

 

  1. The Children’s Butterfly Site. Mid-continent Ecological Science Center, U.S. Geological

Science Center, Ft. Collins, Colo.

 

  1. Status and trends of the Nation’s biological resources. 2 vols. U.S. Department of Interior, U.S. Geological Survey, Reston, Va. 964 pp. (with Mac, M., C. Puckett Haecker, and P. Doran, co-editors).

 

  1. California. Vol. 2, pp. 593-644 in: M. Mac, P. Opler, C. Puckett Haecker, and P. Doran,

co-editors). Status and trends of the nation’s biological resources. U.S. Department of Interior,

U.S. Geological Survey, Reston, Va. 964 pp. (co-authored with S.D. Viers).

 

  1. Common and Scientific names Appendix. Vol. 2, pp. 869-894 in: M. Mac, P. Opler, C.

Puckett Haecker, and P. Doran, co-editors). Status and trends of the nation’s biological

resources. U.S. Department of Interior, U.S. Geological Survey, Reston, Va. 964 pp.

 

  1. Fourth of July butterfly counts (1997). North American Butterfly Association, Baraboo,

Wisc. (Co-edited with Ann B. Swengel).

 

  1. Field Guide to Butterflies of Eastern North America. Revised format. Roger Tory

Peterson Series. Houghton-Mifflin Co., Boston. 486 pp.

 

  1. Systematic Compendium of Arthropods and Annelid Worms of Old-growth and Late

Successional Conifer Forests, Mature Riparian Woods, and of Coarse Woody Debris

Associated Arthropods within the Range of the Northern Spotted Owl (Strix occidentalis

caurina). (co-compiler, Jack D. Lattin.) 1998. National Biological Survey, Midcontinent

Ecological Science Center online publication. [http://webmesc.mesc.nbs.gov/pubs/online/arthropods/systematic_compendium.html]

  1. Field Guide to Butterflies of Western North America. Roger Tory Peterson Series.

Houghton-Mifflin Co., Boston. 540 pp.

 

  1. A new Chlosyne checkerspot from northeast Mexico. (Lepidoptera: Nymphalidae:

Nymphalinae). Holarctic Lepidoptera. 6(1): 23-24.

 

  1. An unusual new Napaea metalmark from Sonora, Mexico. (Lepidoptera: Riodinidae).

Holarctic Lepidoptera 6(1): 25-29. (co-authored with A.D. Warren).

 

  1. A guide to the common insects of Canyonlands National Park. Co-authored with B.C. Ko nratieff, M.J. Weissmann, and P.A. Opler, 16 pages + illustrations, draft.

 

  1. Insects and Related Arthropods of North America. (Coordinator). Northern Prairie

Wildlife Research Center Online, U.S. Geological Survey, Jamestown, ND:.

http://www.npwrc.usgs.gov/resource/insects/insect/index.htm (Version 26JUN2002).

 

  1. Preface, p. vi in: C.V. Covell, Jr. The Butterflies and Moths (Lepidoptera) of Kentucky.

Kentucky State Nature Preserves Commission. Scientific and Technical Series Number 6,

Frankfort, Ky.

 

  1. Revised Recovery Plan for Oregon Silverspot Butterfly. U.S. Fish and Wildlife Service,

Portland, Ore. Co-authored with recovery team.

 

  1. Rapid assessment of butterfly diversity in a montane landscape. Biodivers Conserv

10:1369–1386. (Simonson, S.E., Opler P.A., Stohlgren T.J., Chong G.W., authors).

 

  1. Scientific names list of the butterflies (Papilionoidea and Hesperioidea) of North America

north of Mexico. Butterflies of North America. 2. Contributions of the Gillette Museum of

Arthropod Diversity, Colorado State University, Fort Collins, Colo. 83 pp.

 

  1. Survey of selected insects of Fort Sill, Comanche County, Oklahoma. Report to U.S.

Army (with B.C. Kondratieff, J. Schmidt, and E. Buckner-Opler).

 

  1. Three new Schinia from western North America (Lepidoptera: Noctuidae: Heliothinae).

Holarctic Lepidoptera.

 

  1. Book review: Systematics of western North American Butterflies, T.C. Emmel. Editor.

Journal of the Lepidopterists’ Society.

 

  1. Insects and Diseases of Woody Plants of the Central Rockies (Paperback) 2003

~ Whitney Cranshaw (Author), David Leatherman (Author), William Jacobi 2003. Insects and

Diseases of Woody Plants of the Central Rockies. Colorado State University, Cooperative

Extension, Bulletin 506A, 283 pp. (W. Cranshaw, senior author, also with B. Kondraitieff , D.

Leatherman, B. Jacobi, and L. Mannix).

 

  1. Fixation of type locality for Lycaena acmon Westwood and characterization of the species

and its distribution. The Taxonomic Report 4: 1-5.

 

  1. Scientific names list for butterfly species of North America north of Mexico. Building a global butterfly information system on the Internet: issues and challenge. Staatsmuseum fĂĽr Naturkunde, Stuttgart, Germany, 2 pages (P. Opler and A. Warren).

 

  1. Butterflies of Wyoming: National Park Service Units, Devil’s Tower National Monument,

Crook County and Fort Laramie National Historical Park, Goshen County, 9 pages [Contract

Report].

 

  1. Insects of western North America. 3. Survey of selected insect taxa of Fort Sill, Comanche

County, Oklahoma—Dragonflies (Odonata), Stoneflies (Plecoptera) and selected Moths

(Lepidoptera). Contributions of the C.P. Gillette Museum of Arthropod Diversity, 92 pp.

(March 2004) [Contract report] (with Kondratieff, B.C., J. Schmidt, and M.C. Garhart.).

 

  1. Habitats for North American Butterflies. Holarctic Lepidoptera (with Krizek, G.O.).

 

  1. Douglas C. Ferguson and Paul A. Opler. 2006. Checklist of North American Arctiidae

(Lepidoptera). Zootaxa 1299: 1-33.

 

  1. Larval host-plant records of Asteraceae root-feeding Eucosmini in California and

adjacent states (Tortricidae). Journal of the Lepidopterists’ Society. 60: 189-193. (senior author

Powell, J.A.) .

 

  1. Remembering Don MacNeill. Journal of the Lepidopterists’ Society 60(2): 107-114. (with

Collins, M.M., J.A. Powell, and J.M. Burns).

 

  1. Insects of western North America. 5. Survey of selected insect taxa of Fort Sill, Comanche

County, Oklahoma. Part 1. Selected Coleoptera, Hymenoptera, Lepidoptera, and Orthoptera.

Contributions of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University,

Ft. Collins, 123 pp. (with Kondratieff, B.C., P.A. Opler, J.P. Schmidt, and E.M. Buckner-

Opler).

 

  1. Designation of Lectotype and synonymy for Euchloe lotta BeutenmĂĽller (Lepidoptera:

Pieridae). Zootaxa 1767: 67-68 (12 May 2008).

 

  1. Revised checklist of the tiger moths (Arctiidae, Lepidoptera) of the Continental United.

States and Canada. Zootaxa 1677: 1-23. (first author B.C. Schmidt).

 

  1. Audubon Butterfly DVD: Essentials for Beginners and Gardeners (co-authored with

James A. Ebner). Mastervision, New York, N.Y. 10028.

 

  1. Audubon Videoguide to Butterflies: Common and Endangered. DVD (co-authored with

James A. Ebner). Mastervision, New York, N.Y. 10028.

 

  1. Butterflies and Moths of North America Northern Prairie Wildlife Research Center, U.S.

Geological Survey, Jamestown, North Dakota [www.butterfliesandmoths.org] (P.A. Opler, H.

Pavulaan, and R.E. Stanford, coordinators.Web site).

 

  1. Moths of western North America. University of California Press, Berkeley and Los

Angeles (with J.A. Powell).

 

  1. Opler, Paul A., Kelly Lotts, and Thomas Naberhaus. 2009. The Children’s Butterfly Site.

Bozeman, MT: Big Sky Institute. http://www.kidsbutterfly.org/ (Version 11-21-2009).

 

  1. The high-altitude member, Plebejus cotundra; p. 116; Euphilotes stanfordorum and

Euphilotes ancilla barnesi, descriptions, pp. 125-128 in: Fisher,M.S. The Butterflies of

Colorado. Part 4. Riodinidae and Lycaenidae. Lepidoptera of North America. 7. Contributions

of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University, Ft. Collins. (with A. Warren & M. Fisher). 

 

  1. Genetic, phonetic, and distributional relationships of Nearctic Euchloe (Pieridae,

Pierinae, Anthocharidini). Journal of the Lepidopterists’ Society 65(1): 1-14 (with Werner

Back and Michael A. Miller).

 

  1. The Complete Book of North American Butterflies. Thunder Bay Press. San Diego,

Calif., 192 pp. (editor).

 

  1. From Ancient Capital to South of the Clouds: Butterfly Studies in Wild China, May.
  2. Association for Tropical Lepidoptera, Notes June 2014. (R.M. Pyle, P.A. Opler, J. Chu,
  3. Buckner-Opler, A. Hurst).

 

  1. Searching for western blue butterflies: the case of the Owens River Drainage in

California-Abstract. Program and abstracts, 25th Annual Meeting of the High Country

Lepidopterists – November 7 and 8. 2014, Denver Museum of Nature and Sciences. Denver

Museum of Nature & Sciences Reports #1, November 7, 2014, p. 5.

 

  1. The Plebejus acmon Species Group (*Lycaenidae: Lycaeninae) in Southern California

and Baja California -Abstract. Program and abstracts, 25th Annual Meeting of the High

Country Lepidopterists – November 7 and 8, 2014, Denver Museum of Nature and Sciences.

Denver Museum of Nature & Sciences Reports #1, November 7, 2014, p. 6.

 

  1. Lepidoptera Bioblitz nets hundreds of addition species. The Midden, Winter 2014, pp. 8-
  2. Great Basin National Park. Baker, Nevada.

 

  1. Evolution of the genetic identity of Lange’s Metalmark, a federally endangered butterfly

(Lepidoptera: Riodinidae). Journal of Chemical Development [ B. Proshek, J. DuPuis, A. Engberg, K. Davenport, P Opler, J Powell, and FAH Sperling].

 

  1. Phylogenetic analysis supports the recognition of Albuna beutenmulleri Skinner as a

species distinct from A. pyramidalis Walker (Lepidoptera: Sesiidae). Journal of the

Lepidopterists’ Society. [Taft, W.H., Cognato, A.I., and Opler, P.A.].

 

  1. When COI barcodes deceive: complete genomes reveal introgression in hairstreaks.

Proceedings of the Royal Society B: Biological Sciences. [Cong, Q, Shen, J., Borek, D., Robbins,

R., Opler, P.A, Grishin, N.V.].

 

  1. LepNet: The Lepidoptera of North America Network. Zootaxa. [Seltmann, K.C., Cobb,

N.S., Gall, L.F., […including Opler, P.A. … Zolnerowich, G.].

 

  1. Drepanoidea, Doidae, and Noctuoidea, Notodontidae (part): Pygaerinae, Notodontinae,

Cerurinae, Phalerinae, Periergosinae, Dudusinae, Hemiceratinae. In: Lafontaine, J.D. et al.

The Moths of America north of Mexico. Fascicle 22.1A, Wedge Entomological Research

Foundation. Washington, D.C., 348 pp. [Miller, J.S., Wagner, D.L., Opler, P.A., and

Lafontaine, J.D.].

 

  1. Current endemic butterfly species of the Californian Region and suggested additions. Denver Museum of Nature and Science Report 16. [P.A. Opler and N.V. Grishin].

 

  1. Changes to North American butterfly names. The Taxonomic Report of the International Lepidoptera Survey 9(1): 1-10. [Zhang, J., Q. Cong, J. Shen, P.A. Opler and N.V. Grishin].

 

  1. Comment (Case 3709) – Comments on the proposed conservation of names for western North American Hesperia comma-group subspecies through designation of neotypes (see BZN 75: 88-121 [Case]; BZN 76: 57-59 [Comments]; BZN 76: 135-138 [P.A. Opler, Ken Davenport].

 

  1. Conserve the eco-evolutionary dynamic, not the subspecies: phenological divergence and gene flow between temporal cohorts of Euphilotes ancilla endemic to southern Nevada. Conservation Genetics, February 2020, issue 1, pp. 1-17. [Thompson, D.B., K. McKelvey, P. van Els, G. Andrews, P. Jacoby-Garrett, M. Glenn, C. Kallstrom, K. Pilgrim, and P.A. Opler].

 

  1. Genomics of a complete butterfly continent. [J. Zhang, Q. Cong, J. Shen, P. Opler and N. Grishin]. bioRxiv BIORXIV/2019/829887.
  2. Genomic evidence suggests further changes of butterfly names. The Taxonomic Report of the International Lepidoptera Survey 8(7): 1-40. [Zhang, Cong, Shen, Opler, and Grishin].
  3. A DNA barcode library for the butterflies of North America. PeerJ. Article 11157 [J. D’Ercole, V. Dinca, P.A. Opler, N. Kondla, C. Schmidt, J.D. Phillips, R. Robbins, J.M. Burns, S.E. Miller, N. Grishin, E.V. Zakharov, J.R. DeWaard, S. Ratnasingham and P.D.N. Hebert]. https://peerj.com/articles/11157

 

  1. Genomics-guided refinement of butterfly taxonomy. The Taxonomic Report of the International Lepidoptera Survey 9(3): 1-54. [Zhang, Cong, Shen, Opler, and Grishin].
  2. Noctuoidea, Notodontidae (part): In: Lafontaine, J.D. et al. The Moths of America north of Mexico. Fascicle 22.1B, Wedge Entomological Research Foundation. Washington, D.C., [Miller, J.S., Wagner, D.L., Opler, P.A., and Lafontaine, J.D.].
  3. Gluphisia richersi, new species. Addendum 1. In: Noctuoidea, Notodontidae (part): In: Lafontaine, J.D. et al. The Moths of America north of Mexico. Fascicle 22.1B, Wedge Entomological Research Foundation. Co-authored with C.D. Ferris.
  4. Invited book review: The disappearance of butterflies by Josef H. Reichholf, Polity Press, Cambridge, U.K. and Medford, MA. News of the Lepidopterists’ Society, 63(1): 21.

 

  1. Metamorphosis: George O. Krizek (1932-2021). News of the Lepidopterists’ Society, 63(1): 37.

 

  1. Preface. Butterflies of Joshua Tree National Monument by Gordon Pratt.
  2. Metamorphosis: Ray E. Stanford (1939-2022). News of the Lepidopterists’ Society, 64: —-.
  3. DNA barcodes reveal different speciation scenarios in the four North American Anthocharis Boisduval, Rambur, Duménil & Graslin, [1833] (Lepidoptera: Pieridae: Pierinae: Anthocharidini) species groups. Zootaxa 5194 (4): 519-539. [Paul A. Opler, Todd L. Stout, Werner Back, Jing Zhang, Qian Cong, Jinhui Shen, & Nick Grishin].
  4. Taxonomic discoveries enabled by genomic analysis of butterflies. The Taxonomic Report of the International Lepidoptera Survey 10(7): 1-60. [Zhang, Cong, Shen, Song, Gott, Boyer, Guppy, Kohler, Lamas, Opler, and Grishin].
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